John is a Managing Director and Partner in the Boston office of BCG. He combines strategic acumen and deep scientific expertise to help industry-leading biopharma clients tackle their most critical strategic, operational, and organizational challenges, with a focus on R&D topics. John is a molecular biologist / biophysicist by training, with a PhD in Engineering and Physical Biology from Harvard, and a BS in Biology from MIT. He has several first author publications in leading journals such as Cell and PNAS on topics of genome organization, epigenomic regulation of gene expression, and chromosome stability.